Taxonomy - the study of classification - is a constantly-evolving field. Changes to species lists are regularly proposed as new physical and genetic evidence becomes available, and as different species concepts (which define what should be considered separate species) are explored. These changes regularly occur on a variety of taxonomic scales, but there is little standardization for their acceptance within the wider scientific community. This makes answering the question "How many ungulate species are there?" a challenge to answer, since there is no single response.
Four recent ungulate classifications are presented and compared below, with species grouped based on the most conservative taxonomy. The species lists range from approximately 250 species in conservative taxonomies such as Mammal Species of the World (Wilson & Reeder, 2005) and the IUCN Red List (2024) to over 450 species in Ungulate Taxonomy (Groves and Grubb, 2010). As of 2024, UltimateUngulate.com follows the taxonomy synthesized in the American Society of Mammalogist's online Mammal Diversity Database.
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Ungulates of the World |
What is an UNGULATE? |
Explore the Family Tree |
Mammal Species of the World (Wilson & Reeder, 2005) |
Ungulate Taxonomy (Groves and Grubb, 2011) |
2024 IUCN taxonomy (IUCN Red List, 2024) |
Mammal Diversity Database (ASM, 2024) |
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Order Perissodactyla |
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Family Equidae |
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Equus |
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Equus asinus | Equus africanus | Equus africanus | Equus africanus Equus asinus |
Equus burchellii Equus quagga |
Equus quagga | Equus quagga | Equus quagga |
Equus caballus | Equus ferus Equus przewalskii |
Equus ferus | Equus ferus |
Equus grevyi | Equus grevyi | Equus grevyi | Equus grevyi |
Equus hemionus | Equus hemionus Equus hemippus Equus khur |
Equus hemionus | Equus hemionus |
Equus kiang | Equus kiang | Equus kiang | Equus kiang |
Equus zebra | Equus hartmannae Equus zebra |
Equus zebra | Equus zebra |
Family Tapiridae |
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Tapirus |
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Tapirus bairdii | Tapirella bairdii | Tapirus bairdii | Tapirus bairdii |
Tapirus indicus | Acrocodia indica | Tapirus indicus | Tapirus indicus |
Tapirus pinchaque | Tapirus pinchaque | Tapirus pinchaque | Tapirus pinchaque |
Tapirus terrestris | Tapirus terrestris | Tapirus terrestris | Tapirus terrestris |
Family Rhinocerotidae |
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Ceratotherium |
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Ceratotherium simum | Ceratotherium simum Ceratotherium cottoni |
Ceratotherium simum | Ceratotherium simum |
Dicerorhinus |
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Dicerorhinus sumatrensis | Dicerorhinus sumatrensis | Dicerorhinus sumatrensis | Dicerorhinus sumatrensis |
Diceros |
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Diceros bicornis | Diceros bicornis | Diceros bicornis | Diceros bicornis |
Rhinoceros |
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Rhinoceros sondaicus | Rhinoceros sondaicus | Rhinoceros sondaicus | Rhinoceros sondaicus |
Rhinoceros unicornis | Rhinoceros unicornis | Rhinoceros unicornis | Rhinoceros unicornis |
Order Artiodactyla |
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Family Camelidae |
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Tribe Camelini |
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Camelus |
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Camelus bactrianus | Camelus ferus | Camelus ferus | Camelus bactrianus Camelus ferus |
Camelus dromedarius | - | - | Camelus dromedarius |
Tribe Lamini |
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Lama |
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Lama glama | Lama guanicoe | Lama guanicoe | Lama glama Lama guanicoe |
Vicugna vicugna | Lama mensalis Lama vicugna |
Vicugna vicugna | Lama pacos Lama vicugna |
Family Suidae |
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Babyrousa |
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Babyrousa babyrussa | Babyrousa babyrussa | Babyrousa babyrussa | Babyrousa babyrussa |
Babyrousa bolabatuensis | - | - | - |
Babyrousa celebensis | Babyrousa celebensis | Babyrousa celebensis | Babyrousa celebensis |
Babyrousa togeanensis | Babyrousa togeanensis | Babyrousa togeanensis | Babyrousa togeanensis |
Hylochoerus |
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Hylochoerus meinertzhageni | Hylochoerus ivoriensis Hylochoerus meinertzhageni Hylochoerus rimator |
Hylochoerus meinertzhageni | Hylochoerus meinertzhageni |
Phacochoerus |
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Phacochoerus aethiopicus | Phacochoerus aethiopicus | Phacochoerus aethiopicus | Phacochoerus aethiopicus |
Phacochoerus africanus | Phacochoerus africanus | Phacochoerus africanus | Phacochoerus africanus |
Porcula |
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Sus salvanius | Porcula salvania | Porcula salvania | Porcula salvania |
Potamochoerus |
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Potamochoerus larvatus | Potamochoerus larvatus | Potamochoerus larvatus | Potamochoerus larvatus |
Potamochoerus porcus | Potamochoerus porcus | Potamochoerus porcus | Potamochoerus porcus |
Sus |
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Sus ahoenobarbus | Sus ahoenobarbus | Sus ahoenobarbus | Sus ahoenobarbus |
Sus barbatus | Sus barbatus | Sus barbatus | Sus barbatus |
Sus bucculentus | - | Sus bucculentus | - |
Sus cebifrons | Sus cebifrons | Sus cebifrons | Sus cebifrons |
Sus celebensis | Sus celebensis | Sus celebensis | Sus celebensis |
Sus oliveri | Sus oliveri | Sus oliveri | Sus oliveri |
Sus philippensis | Sus philippensis | Sus philippensis | Sus philippensis |
Sus scrofa | Sus chirodontus Sus cristatus Sus davidi Sus leucomystax Sus moupinensis Sus scrofa Sus taevanys Sus ussuricus Sus vittatus ?Sus meridionalis ?Sus nigripes ?Sus riukiuanus ?Sus sibiricus |
Sus scrofa | Sus domesticus |
Sus verrucosus | Sus blouchi Sus verrucosus |
Sus verrucosus | Sus verrucosus |
Family Tayassuidae |
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Dicotyles |
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Pecari tajacu | Pecari angulatus Pecari crassus Pecari tajacu |
Pecari tajacu | Dicotyles tajacu |
Parachoerus |
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Catagonus wagneri | Catagonus wagneri | Catagonus wagneri | Parachoerus wagneri |
Tayassu |
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Tayassu pecari | Tayassu pecari | Tayassu pecari | Tayassu pecari | Family Hippopotamidae |
Choeropsis |
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Choeropsis liberiensis | Choeropsis heslopi Choeropsis liberiensis |
Choeropsis liberiensis | Choeropsis liberiensis |
Hippopotamus |
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Hippopotamus amphibius | Hippopotamus amphibius | Hippopotamus amphibius | Hippopotamus amphibius |
Suborder Ruminantia |
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Family Tragulidae |
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Hyemoschus |
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Hyemoschus aquaticus | Hyemoschus aquaticus | Hyemoschus aquaticus | Hyemoschus aquaticus |
Moschiola |
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Moschiola meminna | Moschiola indica Moschiola kathygre Moschiola meminna |
Moschiola indica Moschiola kathygre Moschiola meminna |
Moschiola indica Moschiola kathygre Moschiola meminna |
Tragulus |
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Tragulus javanicus | Tragulus javanicus | Tragulus javanicus | Tragulus javanicus |
Tragulus kanchil | Tragulus kanchil | Tragulus kanchil | Tragulus kanchil |
Tragulus napu | Tragulus napu | Tragulus napu | Tragulus napu |
Tragulus nigricans | Tragulus nigricans | Tragulus nigricans | Tragulus nigricans |
Tragulus versicolor | Tragulus versicolor | Tragulus versicolor | Tragulus versicolor |
Tragulus williamsoni | Tragulus williamsoni | Tragulus williamsoni | Tragulus williamsoni |
Family Giraffidae |
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Giraffa |
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Giraffa camelopardalis | Giraffa angolensis Giraffa antiquorum Giraffa camelopardalis Giraffa giraffa Giraffa peralta Giraffa reticulata Giraffa thornicrofti Giraffa tippelskirchi |
Giraffa camelopardalis | Giraffa camelopardalis Giraffa giraffa Giraffa reticulata Giraffa tippelskirchi |
Okapia |
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Okapia johnstoni | Okapia johnstoni | Okapia johnstoni | Okapia johnstoni |
Family Antilocapridae |
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Antilocapra americana | Antilocapra americana | Antilocapra americana | Antilocapra americana |
Family Moschidae |
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Moschus anhuiensis | Moschus anhuiensis | Moschus anhuiensis | Moschus anhuinensis |
Moschus berezovskii | Moschus berezovskii | Moschus berezovskii | Moschus berezovskii |
Moschus chrysogaster | Moschus chrysogaster | Moschus chrysogaster | Moschus chrysogaster |
Moschus cupreus | Moschus cupreus | Moschus cupreus | Moschus cupreus |
Moschus fuscus | Moschus fuscus | Moschus fuscus | Moschus fuscus |
Moschus leucogaster | Moschus leucogaster | Moschus leucogaster | Moschus leucogaster |
Moschus moschiferus | Moschus moschiferus | Moschus moschiferus | Moschus moschiferus |
Family Cervidae |
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Subfamily Capreolinae |
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Tribe Alceini |
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Alces |
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Alces alces Alces americanus |
Alces alces Alces americanus |
Alces alces | Alces alces |
Tribe Capreolini |
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Capreolus |
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Capreolus capreolus | Capreolus capreolus | Capreolus capreolus | Capreolus capreolus |
Capreolus pygargus | Capreolus pygargus | Capreolus pygargus | Capreolus pygargus |
Hydropotes |
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Hydropotes inermis | Hydropotes inermis | Hydropotes inermis | Hydropotes inermis |
Tribe Odocoileini |
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Blastocerus |
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Blastocerus dichotomus | Blastocerus dichotomus | Blastocerus dichotomus | Blastocerus dichotomus |
Hippocamelus |
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Hippocamelus antisensis | Hippocamelus antisensis | Hippocamelus antisensis | Hippocamelus antisensis |
Hippocamelus bisulcus | Hippocamelus bisulcus | Hippocamelus bisulcus | Hippocamelus bisulcus |
Mazama |
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Mazama americana Mazama bororo |
Mazama americana Mazama bororo Mazama gualea Mazama jucunda Mazama trinitatis Mazama whitelyi Mazama zamora Mazama zetta |
Mazama americana Mazama bororo |
Mazama americana Mazama jucunda Mazama rufa |
Mazama bricenii Mazama rufina |
Mazama bricenii Mazama rufina |
Mazama bricenii Mazama rufina |
Mazama rufina |
Mazama chunyi | Mazama chunyi | Mazama chunyi | Mazama chunyi |
Mazama gouazoubira | Mazama cita |
Mazama gouazoubira Mazama nemorivaga |
Bisbalus citus Passalites nemorivagus Subulo gouazoubira |
Mazama nana | Mazama nana | Mazama nana | Mazama nanus |
Mazama temama | Mazama temama | Mazama temama | Mazama temama |
Odocoileus |
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Odocoileus hemionus | Odocoileus hemionus | Odocoileus hemionus | Odocoileus hemionus |
Mazama pandora | Mazama pandora | Mazama pandora | Odocoileus pandora |
Odocoileus virginianus | Odocoileus virginianus | Odocoileus virginianus | Odocoileus virginianus |
(With a note that the genus Odocoileus requires significant taxonomic revision) | |||
Ozotoceros |
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Ozotoceros bezoarticus | Ozotoceros bezoarticus | Ozotoceros bezoarticus | Ozotoceros bezoarticus |
Pudu |
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Pudu mephistophiles | Pudu mephistophiles | Pudu mephistophiles | (Pudella carlae) Pudella mephistophila |
Pudu puda | Pudu puda | Pudu puda | Pudu puda |
Rangifer |
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Rangifer tarandus | Rangifer tarandus | Rangifer tarandus | Rangifer tarandus |
Subfamily Cervinae |
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Tribe Cervini |
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Axis |
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Axis axis | Axis axis | Axis axis | Axis axis |
Axis calamianensis | Axis calamianensis | Axis calamianensis | Axis calamianensis |
Axis kuhlii | Axis kuhlii | Axis kuhlii | Axis kuhlii |
Axis porcinus | Axis annamiticus Axis porcinus |
Axis porcinus | Axis porcinus |
Cervus |
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Cervus elaphus | Cervus alashanicus Cervus bactrianus Cervus canadensis Cervus corsicanus Cervus elaphus Cervus hanglu Cervus macneilli Cervus maral Cervus pannoniensis Cervus wallichii Cervus xanthopygus Cervus yarkandensis |
Cervus canadensis Cervus elaphus Cervus hanglu |
Cervus canadensis Cervus elaphus Cervus hanglu |
Cervus nippon | Cervus aplodontus Cervus hortulorum Cervus nippon Cervus pseudaxis Cervus pulchellus Cervus sichuanicus Cervus taiouanus |
Cervus nippon | Cervus nippon |
Przewalskium albirostris | Cervus albirostris | Cervus albirostris | Cervus albirostris |
Dama |
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Dama dama | Dama dama Dama mesopotamica |
Dama dama Dama mesopotamica |
Dama dama Dama mesopotamica |
Elaphurus |
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Elaphurus davidianus | Elaphurus davidianus | Elaphurus davidianus | Elaphurus davidianus |
Rucervus |
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Rucervus duvaucelii | Rucervus branderi Rucervus duvaucelii Rucervis ranjitsinhi |
Rucervus duvaucelii | Rucervus duvaucelii |
Rucervus elsii | Panolia eldii Panolia siamensis Panolia thamin |
Rucervus eldii | Rucervus eldii |
Rucervus schomburgki | Rucervus schomburgki | Rucervus schomburgki | Rucervus schomburgki |
Rusa |
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Rusa alfredi | Cervus alfredi | Rusa alfredi | Rusa alfredi |
Rusa marianna | Cervus barandanus Cervus mariannus Cervus nigellus |
Rusa marianna | Rusa marianna |
Rusa timorensis | Cervus timorensis | Rusa timorensis | Rusa timorensis |
Rusa unicolor | Cervus equinus Cervus unicolor |
Rusa unicolor | Rusa unicolor |
Tribe Muntiacini |
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Elaphodus |
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Elaphodus cephalophus | Elaphodus cephalophus | Elaphodus cephalophus | Elaphodus cephalophus |
Muntiacus |
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Muntiacus atherodes | Muntiacus atherodes | Muntiacus atherodes | Muntiacus atherodes |
Muntiacus crinifrons | Muntiacus crinifrons | Muntiacus crinifrons | Muntiacus crinifrons |
Muntiacus feae | Muntiacus feae | Muntiacus feae | Muntiacus feae |
Muntiacus gongshanensis | Muntiacus gongshanensis | Muntiacus gongshanensis | Muntiacus gongshanensis |
Muntiacus muntjak | Muntiacus aureus |
Muntiacus montanus Muntiacus muntjak Muntiacus vaginalis |
Muntiacus malabaricus Muntiacus muntjak Muntiacus vaginalis |
Muntiacus puhoatensis | Muntiacus puhoatensis | Muntiacus puhoatensis | Muntiacus puhoatensis |
Muntiacus putaoensis | Muntiacus putaoensis | Muntiacus putaoensis | Muntiacus putaoensis |
Muntiacus reevesi | Muntiacus reevesi | Muntiacus reevesi | Muntiacus reevesi |
Muntiacus rooseveltorum | Muntiacus rooseveltorum | Muntiacus rooseveltorum | Muntiacus rooseveltorum |
Muntiacus truongsonensis | Muntiacus truongsonensis | Muntiacus truongsonensis | Muntiacus truongsonensis |
Muntiacus vuquangensis | Muntiacus vuquangensis | Muntiacus vuquangensis | Muntiacus vuquangensis |
Family Bovidae |
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Tribe Aepycerotini |
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Aepyceros |
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Aepyceros melampus | Aepyceros melampus Aepyceros petersi |
Aepyceros melampus | Aepyceros melampus |
Tribe Alcelaphini |
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Alcelaphus |
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Alcelaphus buselaphus Alcelaphus caama Alcelaphus lichtensteini |
Alcelaphus buselaphus Alcelaphus caama Alcelaphus cokii Alcelaphus lelwel Alcelaphus lichtensteini Alcelaphus major Alcelaphus swaynei Alcelaphus tora |
Alcelaphus buselaphus | Alcelaphus buselaphus |
Beatragus |
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Beatragus hunteri | Beatragus hunteri | Beatragus hunteri | Beatragus hunteri |
Connochaetes |
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Connochaetes gnou | Connochaetes gnou | Connochaetes gnou | Connochetes gnou |
Connochaetes taurinus | Connochaetes albojubatus Connochaetes johnstoni Connochaetes mearnsi Connochaetes taurinus |
Connochaetes taurinus | Connochaetes taurinus |
Damaliscus |
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Damaliscus korrigum Damaliscus lunatus Damaliscus superstes |
Damaliscus eurus Damaliscus jimela Damaliscus korrigum Damaliscus lunatus Damaliscus selousi Damaliscus superstes Damaliscus tiang Damaliscus topi Damaliscus ugandae |
Damaliscus lunatus | Damaliscus lunatus |
Damaliscus pygargus | Damaliscus phillipsi Damaliscus pygargus |
Damaliscus pygargus | Damaliscus pygargus |
Tribe Antilopini |
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Ammodorcas |
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Ammodorcas clarkei | Ammodorcas clarkei | Ammodorcas clarkei | Ammodorcas clarkei |
Antidorcas |
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Antidorcas marsupialis | Antidorcas angolensis Antidorcas hofmeyri Antidorcas marsupialis |
Antidorcas marsupialis | Antidorcas marsupialis |
Antilope |
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Antilope cervicapra | Antilope cervicapra | Antilope cervicapra | Antilope cervicapra |
Dorcatragus |
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Dorcatragus megalotis | Dorcatragus megalotis | Dorcatragus megalotis | Dorcatragus megalotis |
Eudorcas |
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Eudorcas rufifrons | Eudorcas albonotata Eudorcas rufifrons Eudorcas tilonura |
Eudorcas albonotata Eudorcas rufifrons Eudorcas tilonura |
Eudorcas albonotata Eudorcas rufifrons Eudorcas tilonura |
Eudorcas rufina | Eudorcas rufina | - | Eudorcas rufina |
Eudorcas thomsonii | Eudorcas nasalis Eudorcas thomsoni |
Eudorcas thomsonii | Eudorcas thomsonii |
Gazella |
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Gazella arabica Gazella erlangeri |
Gazella acaciae Gazella arabica Gazella bilkis Gazella cora Gazella dareshurii Gazella erlangeri Gazella muscatensis |
Gazella arabica Gazella bilkis |
Gazella arabica |
Gazella bennettii | Gazella bennettii Gazella christyi Gazella fuscifrons Gazella shikarii Gazella salinarum |
Gazella bennettii | Gazella bennettii |
Gazella cuvieri | Gazella cuvieri | Gazella cuvieri | Gazella cuvieri |
Gazella dorcas Gazella saudiya |
Gazella dorcas Gazella pelzelni Gazella saudiya |
Gazella dorcas Gazella saudiya |
Gazella dorcas |
Gazella gazella |
Gazella gazella Gazella karamii |
Gazella gazella | Gazella gazella |
Gazella leptoceros | Gazella leptoceros | Gazella leptoceros | Gazella leptoceros |
Gazella spekei | Gazella spekei | Gazella spekei | Gazella spekei |
Gazella subgutturosa | Gazella gracilicornis Gazella marica Gazella subgutturosa Gazella yarkandensis |
Gazella marica Gazella subgutturosa |
Gazella marica Gazella subgutturosa |
Litocranius |
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Litocranius walleri | Litocranius sclateri Litocranius walleri |
Litocranius walleri | Litocranius walleri |
Madoqua |
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Madoqua guentheri | Madoqua guentheri Madoqua smithii |
Madoqua guentheri | Madoqua guentheri |
Madoqua kirkii | Madoqua cavendishi Madoqua damarensis Madoqua hindei Madoqua kirkii Madoqua thomasi |
Madoqua kirkii | Madoqua cavendishi Madoqua damarensis Madoqua kirkii Madoqua thomasi |
Madoqua piacentinii | Madoqua piacentinii | Madoqua piacentinii | Madoqua piacentinii |
Madoqua saltiana | Madoqua hararensis Madoqua lawrancei Madoqua phillipsi Madoqua saltiana Madoqua swaynei |
Madoqua saltiana | Madoqua saltiana |
Nanger |
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Nanger dama | Nanger dama | Nanger dama | Nanger dama |
Nanger granti | Nanger granti Nanger notata Nanger petersi |
Nanger granti | Nanger granti Nanger notatus Nanger petersii |
Nanger soemmerringii | Nanger soemmerringi | Nanger soemmerringii | Nanger soemmerringii |
Ourebia |
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Ourebia ourebi | Ourebia hastata Ourebia montana Ourebia ourebi Ourebia quadriscopa |
Ourebia ourebi | Ourebia ourebi |
Procapra |
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Procapra gutturosa | Procapra gutturosa | Procapra gutturosa | Procapra gutturosa |
Procapra picticaudata | Procapra picticaudata | Procapra picticaudata | Procapra picticaudata |
Procapra przewalskii | Procapra przewalskii | Procapra przewalskii | Procapra przewalskii |
Raphicerus |
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Raphicerus campestris | Raphicerus campestris | Raphicerus campestris | Raphicerus campestris |
Raphicerus melanotis | Raphicerus melanotis | Raphicerus melanotis | Raphicerus melanotis |
Raphicerus sharpei | Raphicerus colonicus Raphicerus sharpei |
Raphicerus sharpei | Raphicerus sharpei |
Saiga |
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Saiga tatarica | Saiga mongolica Saiga tartarica |
Saiga tatarica | Saiga tatarica |
Tribe Caprini |
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Ammotragus |
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Ammotragus lervia | Ammotragus lervia | Ammotragus lervia | Ammotragus lervia |
Arabitragus |
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Hemitragus jayakari | Arabitragus jayakari | Arabitragus jayakari | Arabitragus jayakari |
Budorcas |
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Budrocas taxicolor | Budorcas bedfordi Budorcas taxicolor Budorcas tibetana Budorcas whitei |
Budorcas taxicolor | Budorcas taxicolor Budorcas tibetana |
Capra |
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Capra caucasica | Capra caucasica Capra cylindricornis |
Capra caucasica Capra cylindricornis |
Capra caucasica Capra cylindricornis |
Capra falconeri | Capra falconeri | Capra falconeri | Capra falconeri |
Capra hircus | Capra aegagrus | Capra aegagrus | Capra aegagrus Capra hircus |
Capra ibex | Capra ibex | Capra ibex | Capra ibex |
Capra nubiana | Capra nubiana | Capra nubiana | Capra nubiana |
Capra pyrenaica | Capra pyrenaica | Capra pyrenaica | Capra pyrenaica |
Capra sibirica | Capra sibirica | Capra sibirica | Capra sibirica |
Capra walie | Capra walie | Capra walie | Capra walie |
Capricornis |
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Capricornis crispus | Capricornis crispus | Capricornis crispus | Capricornis crispus |
Capricornis milneedwardsi Capricornis sumatrensis Capricornis thar |
Capricornis maritimus |
Capricornis sumatrensis | Capricornis sumatrensis |
Capricornis rubidus | Capricornis rubidus | Capricornis rubidus | Capricornis rubidus |
Capricornis swinhoei | Capricornis swinhoii | Capricornis swinhoei | Capricornis swinhoei |
Hemitragus |
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Hemitragus jemlahicus | Hemitragus jemlahicus | Hemitragus jemlahicus | Hemitragus jemlahicus |
Naemorhedus |
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Naemorhedus baileyi | Nemorhaedus baileyi | Naemorhedus baileyi | Naemorhedus baileyi Naemorhedus cranbrooki |
Naemorhedus caudatus | Nemorhaedus caudatus | Naemorhedus caudatus | Naemorhedus caudatus |
Naemorhedus goral | Nemorhaedus bedfordi Nemorhaedus goral |
Naemorhedus goral | Naemorhedus goral |
Naemorhedus griseus | Nemorhaedus evansi Nemorhaedus griseus |
(Naemorhedus griseus) | Naemorhedus evansi Naemorhedus griseus |
Nilgiritragus |
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Nilgiritragus hylocrius | Nilgiritragus hylocrius | Nilgiritragus hylocrius | Nilgiritragus hylocrius |
Oreamnos |
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Oreamnos americanus | Oreamnos americanus | Oreamnos americanus | Oreamnos americanus |
Ovibos |
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Ovibos moschatus | Ovibos moschatus | Ovibos moschatus | Ovibos moschatus |
Ovis |
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Ovis ammon | Ovis ammon Ovis collium Ovis darwini Ovis hodgsoni Ovis jubata Ovis karelini Ovis nigrimontana Ovis polii |
Ovis ammon | Ovis ammon |
Ovis aries | Ovis arabica Ovis bochariensis Ovis cycloceros Ovis gmelini Ovis isphaganica Ovis laristanica Ovis punjabensis Ovis severtzovi Ovis vignei |
Ovis gmelini Ovis vignei |
Ovis aries Ovis gmelinii Ovis vignei |
Ovis canadensis | Ovis canadensis | Ovis canadensis | Ovis canadensis |
Ovis dalli | Ovis dalli | Ovis dalli | Ovis dalli |
Ovis nivicola | Ovis nivicola | Ovis nivicola | Ovis nivicola |
Pantholops |
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Pantholops hodgsonii | Pantholops hodgsonii | Pantholops hodgsonii | Pantholops hodgsonii |
Pseudois |
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Pseudois nayaur Pseudois schaeferi |
Pseudois nayaur Pseudois schaeferi |
Pseudois nayaur | Pseudois nayaur |
Rupicapra |
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Rupicapra pyrenaica | Rupicapra ornata Rupicapra parva Rupicapra pyrenaica |
Rupicapra pyrenaica | Rupicapra pyrenaica |
Rupicapra rupicapra | Rupicapra asiatica Rupicapra carpatica Rupicapra rupicapra ?Rupicapra balcanica |
Rupicapra rupicapra | Rupicapra rupicapra |
Tribe Cephalophini |
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Cephalophorus |
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Cephalophus brookei Cephalophus ogilbyi |
Cephalophus brookei Cephalophus crusalbum Cephalophus ogilbyi |
Cephalophus ogilbyi | Cephalophorus brookei Cephalophorus ogilbyi |
Cephalophus callipygus | Cephalophus callipygus | Cephalophus callipygus | Cephalophorus callipygus |
Cephalophus leucogaster | Cephalophus arrhenii Cephalophus leucogaster |
Cephalophus leucogaster | Cephalophorus leucogaster |
Cephalophus natalensis | Cephalophus harveyi Cephalophus natalensis |
Cephalophus harveyi Cephalophus natalensis |
Cephalophorus natalensis |
Cephalophus niger | Cephalophus niger | Cephalophus niger | Cephalophorus niger |
Cephalophus nigrifrons | Cephalophus fosteri Cephalophus hooki Cephalophus hypoxanthus Cephalophus nigrifrons Cephalophus rubidus |
Cephalophus nigrifrons | Cephalophorus kivuensis Cephalophorus nigrifrons Cephalophorus rubidus |
Cephalophus rufilatus | Cephalophus rufilatus | Cephalophus rufilatus | Cephalophorus rufilatus |
Cephalophus weynsi | Cephalophus johnstoni Cephalophus lestradei Cephalophus weynsi |
Cephalophus weynsi | Cephalophorus weynsi |
Cephalophula |
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Cephalophus zebra | Cephalophus zebra | Cephalophus zebra | Cephalophula zebra |
Cephalophus |
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Cephalophus dorsalis | Cephalophus castaneus Cephalophus dorsalis |
Cephalophus dorsalis | Cephalophus dorsalis |
Cephalophus jentinki | Cephalophus jentinki | Cephalophus jentinki | Cephalophus jentinki |
Cephalophus silvicultor | Cephalophus curticeps Cephalophus silvicultor |
Cephalophus silvicultor | Cephalophus silvicultor |
Cephalophus spadix | Cephalophus spadix | Cephalophus spadix | Cephalophus spadix |
Leucocephalophus |
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Cephalophus adersi | Cephalophus adersi | Cephalophus adersi | Leucocephalophus adersi |
Philantomba |
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Philantomba maxwellii | Philantomba maxwelli Philantomba walteri |
Philantomba maxwellii Philantomba walteri |
Philantomba maxwellii Philantomba walteri |
Philantomba monticola | Philantomba aequatorialis Philantomba anchietae Philantomba bicolor Philantomba congicus Philantomba defriesi Philantomba hecki Philantomba lugens Philantomba melanorheus Philantomba monticola Philantomba simpsoni |
Philantomba monticola | Philantomba monticola |
Sylvicapra |
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Sylvicapra grimmia | Sylvicapra coronatus Sylvicapra grimmia Sylvicapra pallidior |
Sylvicapra grimmia | Sylvicapra grimmia |
Tribe Hippotragini |
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Addax |
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Addax nasomaculatus | Addax nasomaculatus | Addax nasomaculatus | Addax nasomaculatus |
Hippotragus |
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Hippotragus equinus | Hippotragus equinus | Hippotragus equinus | Hippotragus equinus |
Hippotragus leucophaeus | Hippotragus leucophaeus | Hippotragus leucophaeus | Hippotragus leucophaeus |
Hippotragus niger | Hippotragus niger Hippotragus roosevelti |
Hippotragus niger | Hippotragus niger |
Oryx |
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Oryx beisa | Oryx beisa Oryx callotis Oryx gallarum |
Oryx beisa | Oryx beisa |
Oryx dammah | Oryx dammah | Oryx dammah | Oryx dammah |
Oryx gazella | Oryx gazella | Oryx gazella | Oryx gazella |
Oryx leucoryx | Oryx leucoryx | Oryx leucoryx | Oryx leucoryx |
Tribe Neotragini |
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Neotragus |
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Neotragus pygmaeus | Neotragus pygmaeus | Neotragus pygmaeus | Neotragus pygmaeus |
Nesotragus |
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Neotragus batesi | Neotragus batesi | Neotragus batesi | Nesotragus batesi |
Neotragus moschatus | Neotragus kirchenpaueri Neotragus livingstonianus |
Nesotragus moschatus | Nesotragus moschatus |
Tribe Oreotragini |
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Oreotragus |
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Oreotragus oreotragus | Oreotragus aceratos Oreotragus aureus Oreotragus centralis Oreotragus oreotragus Oreotragus porteousi Oreotragus saltatrixoides Oreotragus schillingsi Oreotragus somalicus Oreotragus stevensoni Oreotragus transvaalensis Oreotragus tyleri |
Oreotragus oreotragus | Oreotragus oreotragus |
Tribe Reduncini |
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Kobus |
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Kobus ellipsiprymnus | Kobus defassa Kobus ellipsiprymnus |
Kobus ellipsiprymnus | Kobus ellipsiprymnus |
Kobus kob | Kobus kob Kobus leucotis Kobus loderi Kobus thomasi |
Kobus kob | Kobus kob |
Kobus leche | Kobus anselli Kobus kafuensis Kobus leche Kobus robertsi Kobus smithemani |
Kobus leche | Kobus leche |
Kobus megaceros | Kobus megaceros | Kobus megaceros | Kobus megaceros |
Kobus vardonii | Kobus vardoni | Kobus vardonii | Kobus vardonii |
Pelea |
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Pelea capreolus | Pelea capreolus | Pelea capreolus | Pelea capreolus |
Redunca |
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Redunca arundinum | Redunca arundinum Redunca occidentalis |
Redunca arundinum | Redunca arundinum |
Redunca fulvorufula | Redunca adamauae Redunca chanleri Redunca fulvorufula |
Redunca fulvorufula | Redunca fulvorufula |
Redunca redunca | Redunca bohor Redunca cottoni Redunca nigeriensis Redunca redunca |
Redunca redunca | Redunca redunca |
Subfamily Bovinae |
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Tribe Boselaphini |
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Boselaphus |
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Boselaphus tragocamelus | Boselaphus tragocamelus | Boselaphus tragocamelus | Boselaphus tragocamelus |
Tetracerus |
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Tetracerus quadricornis | Tetracerus quadricornis | Tetracerus quadricornis | Tetracerus quadricornis |
Tribe Bovini |
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Bos |
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Bison bison | Bos bison | Bison bison | Bos bison |
Bison bonasus | Bos bonasus Bos caucasicus |
Bison bonasus | Bos bonasus |
Bos frontalis | Bos gaurus | Bos gaurus | Bos frontalis Bos gaurus |
Bos grunniens | Bos mutus | Bos mutus | Bos grunniens Bos mutus |
Bos javanicus | Bos javanicus | Bos javanicus | Bos javanicus |
Bos sauveli | Bos sauveli | Bos sauveli | Bos sauveli |
Bos taurus | Bos primigenius | Bos primigenius | Bos primigenius Bos taurus |
Bubalus |
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Bubalus bubalis | Bubalus arnee | Bubalus arnee | Bubalus arnee Bubalus bubalis |
Bubalus depressicornis | Bubalus depressicornis | Bubalus depressicornis | Bubalus depressicornis |
Bubalus mindorensis | Bubalus mindorensis | Bubalus mindorensis | Bubalus mindorensis |
Bubalus quarlesi | Bubalus quarlesi | Bubalus quarlesi | Bubalus quarlesi |
Pseudoryx |
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Pseudoryx nghetinhensis | Pseudoryx nghetinhensis | Pseudoryx nghetinhensis | Pseudoryx nghetinhensis |
Syncerus |
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Syncerus caffer | Syncerus brachyceros Syncerus caffer Syncerus mathewsi Syncerus nanus |
Syncerus caffer | Syncerus caffer |
Tribe Tragelaphini |
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Tragelaphus |
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Taurotragus derbianus | Taurotragus derbianus | Tragelaphus derbianus | Tragelaphus derbianus |
Taurotragus oryx | Taurotragus oryx | Tragelaphus oryx | Tragelaphus oryx |
Tragelaphus angasii | Nyala angasii | Tragelaphus angasii | Tragelaphus angasii |
Tragelaphus buxtoni | Tragelaphus buxtoni | Tragelaphus buxtoni | Tragelaphus buxtoni |
Tragelaphus eurycerus | Tragelaphus euryceros | Tragelaphus eurycerus | Tragelaphus eurycerus |
Tragelaphus imberbis | Ammelaphus australis Ammelaphus imberbis |
Tragelaphus imberbis | Tragelaphus imberbis |
Tragelaphus scriptus | Tragelaphus bor Tragelaphus decula Tragelaphus fasciatus Tragelaphus meneliki Tragelaphus ornatus Tragelaphus phaleratus Tragelaphus scriptus Tragelaphus sylvaticus |
Tragelaphus scriptus | Tragelaphus scriptus Tragelaphus sylvaticus |
Tragelaphus spekii | Tragelaphus gratus Tragelaphus larkenii Tragelaphus selousi Tragelaphus spekii Tragelaphus sylvestris ?Tragelaphus ugallae |
Tragelaphus spekii | Tragelaphus spekii |
Tragelaphus strepsiceros | Strepsiceros chora Strepsiceros cottoni Strepsiceros strepsiceros Strepsiceros zambesiensis |
Tragelaphus strepsiceros | Tragelaphus strepsiceros |