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Ungulate Taxonomies
A comparison of classifications

Taxonomy - the study of classification - is a constantly-evolving field. Changes to species lists are regularly proposed as new physical and genetic evidence becomes available, and as different species concepts (which define what should be considered separate species) are explored. These changes regularly occur on a variety of taxonomic scales, but there is little standardization for their acceptance within the wider scientific community. This makes answering the question "How many ungulate species are there?" a challenge to answer, since there is no single response.

Four recent ungulate classifications are presented and compared below, with species grouped based on the most conservative taxonomy. The species lists range from approximately 250 species in conservative taxonomies such as Mammal Species of the World (Wilson & Reeder, 2005) and the IUCN Red List (2024) to over 450 species in Ungulate Taxonomy (Groves and Grubb, 2010). As of 2024, UltimateUngulate.com follows the taxonomy synthesized in the American Society of Mammalogist's online Mammal Diversity Database.

Continue below to the taxonomies,
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Ungulates of
 the World
What is an
UNGULATE?
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Family Tree

Mammal Species
of the World

(Wilson & Reeder, 2005)
Ungulate Taxonomy
(Groves and Grubb, 2011)
2024 IUCN taxonomy
(IUCN Red List, 2024)
Mammal Diversity Database
(ASM, 2024)
 
Family Equidae
Equus
Equus asinus Equus africanus Equus africanus Equus africanus
Equus asinus
Equus burchellii
Equus quagga
Equus quagga Equus quagga Equus quagga
Equus caballus Equus ferus
Equus przewalskii
Equus ferus Equus ferus
Equus grevyi Equus grevyi Equus grevyi Equus grevyi
Equus hemionus Equus hemionus
Equus hemippus
Equus khur
Equus hemionus Equus hemionus
Equus kiang Equus kiang Equus kiang Equus kiang
Equus zebra Equus hartmannae
Equus zebra
Equus zebra Equus zebra
 
Family Tapiridae
Tapirus
Tapirus bairdii Tapirella bairdii Tapirus bairdii Tapirus bairdii
Tapirus indicus Acrocodia indica Tapirus indicus Tapirus indicus
Tapirus pinchaque Tapirus pinchaque Tapirus pinchaque Tapirus pinchaque
Tapirus terrestris Tapirus terrestris Tapirus terrestris Tapirus terrestris
 
Ceratotherium
Ceratotherium simum Ceratotherium simum
Ceratotherium cottoni
Ceratotherium simum Ceratotherium simum
Dicerorhinus
Dicerorhinus sumatrensis Dicerorhinus sumatrensis Dicerorhinus sumatrensis Dicerorhinus sumatrensis
Diceros
Diceros bicornis Diceros bicornis Diceros bicornis Diceros bicornis
Rhinoceros
Rhinoceros sondaicus Rhinoceros sondaicus Rhinoceros sondaicus Rhinoceros sondaicus
Rhinoceros unicornis Rhinoceros unicornis Rhinoceros unicornis Rhinoceros unicornis
 
Family Camelidae
Tribe Camelini
Camelus
Camelus bactrianus Camelus ferus Camelus ferus Camelus bactrianus
Camelus ferus
Camelus dromedarius  -  - Camelus dromedarius
 
Tribe Lamini
Lama
Lama glama Lama guanicoe Lama guanicoe Lama glama
Lama guanicoe
Vicugna vicugna Lama mensalis
Lama vicugna
Vicugna vicugna Lama pacos
Lama vicugna
 
Family Suidae
Babyrousa
Babyrousa babyrussa Babyrousa babyrussa Babyrousa babyrussa Babyrousa babyrussa
Babyrousa bolabatuensis  -  -  -
Babyrousa celebensis Babyrousa celebensis Babyrousa celebensis Babyrousa celebensis
Babyrousa togeanensis Babyrousa togeanensis Babyrousa togeanensis Babyrousa togeanensis
Hylochoerus
Hylochoerus meinertzhageni Hylochoerus ivoriensis
Hylochoerus meinertzhageni
Hylochoerus rimator
Hylochoerus meinertzhageni Hylochoerus meinertzhageni
Phacochoerus
Phacochoerus aethiopicus Phacochoerus aethiopicus Phacochoerus aethiopicus Phacochoerus aethiopicus
Phacochoerus africanus Phacochoerus africanus Phacochoerus africanus Phacochoerus africanus
Porcula
Sus salvanius Porcula salvania Porcula salvania Porcula salvania
Potamochoerus
Potamochoerus larvatus Potamochoerus larvatus Potamochoerus larvatus Potamochoerus larvatus
Potamochoerus porcus Potamochoerus porcus Potamochoerus porcus Potamochoerus porcus
Sus
Sus ahoenobarbus Sus ahoenobarbus Sus ahoenobarbus Sus ahoenobarbus
Sus barbatus Sus barbatus Sus barbatus Sus barbatus
Sus bucculentus  - Sus bucculentus  -
Sus cebifrons Sus cebifrons Sus cebifrons Sus cebifrons
Sus celebensis Sus celebensis Sus celebensis Sus celebensis
Sus oliveri Sus oliveri Sus oliveri Sus oliveri
Sus philippensis Sus philippensis Sus philippensis Sus philippensis
Sus scrofa Sus chirodontus
Sus cristatus
Sus davidi
Sus leucomystax
Sus moupinensis
Sus scrofa
Sus taevanys
Sus ussuricus
Sus vittatus
?Sus meridionalis
?Sus nigripes
?Sus riukiuanus
?Sus sibiricus
Sus scrofa

Sus domesticus
Sus scrofa

Sus verrucosus Sus blouchi
Sus verrucosus
Sus verrucosus Sus verrucosus
 
Dicotyles
Pecari tajacu Pecari angulatus
Pecari crassus
Pecari tajacu
Pecari tajacu Dicotyles tajacu
Parachoerus
Catagonus wagneri Catagonus wagneri Catagonus wagneri Parachoerus wagneri
Tayassu
Tayassu pecari Tayassu pecari Tayassu pecari Tayassu pecari
 
Choeropsis
Choeropsis liberiensis Choeropsis heslopi
Choeropsis liberiensis
Choeropsis liberiensis Choeropsis liberiensis
Hippopotamus
Hippopotamus amphibius Hippopotamus amphibius Hippopotamus amphibius Hippopotamus amphibius
 
Suborder Ruminantia
Family Tragulidae
Hyemoschus
Hyemoschus aquaticus Hyemoschus aquaticus Hyemoschus aquaticus Hyemoschus aquaticus
Moschiola
Moschiola meminna Moschiola indica
Moschiola kathygre
Moschiola meminna
Moschiola indica
Moschiola kathygre
Moschiola meminna
Moschiola indica
Moschiola kathygre
Moschiola meminna
Tragulus
Tragulus javanicus Tragulus javanicus Tragulus javanicus Tragulus javanicus
Tragulus kanchil Tragulus kanchil Tragulus kanchil Tragulus kanchil
Tragulus napu Tragulus napu Tragulus napu Tragulus napu
Tragulus nigricans Tragulus nigricans Tragulus nigricans Tragulus nigricans
Tragulus versicolor Tragulus versicolor Tragulus versicolor Tragulus versicolor
Tragulus williamsoni Tragulus williamsoni Tragulus williamsoni Tragulus williamsoni
 
Family Giraffidae
Giraffa
Giraffa camelopardalis Giraffa angolensis
Giraffa antiquorum
Giraffa camelopardalis
Giraffa giraffa
Giraffa peralta
Giraffa reticulata
Giraffa thornicrofti
Giraffa tippelskirchi
Giraffa camelopardalis Giraffa camelopardalis
Giraffa giraffa
Giraffa reticulata
Giraffa tippelskirchi
Okapia
Okapia johnstoni Okapia johnstoni Okapia johnstoni Okapia johnstoni
 
Antilocapra americana Antilocapra americana Antilocapra americana Antilocapra americana
 
Family Moschidae
Moschus anhuiensis Moschus anhuiensis Moschus anhuiensis Moschus anhuinensis
Moschus berezovskii Moschus berezovskii Moschus berezovskii Moschus berezovskii
Moschus chrysogaster Moschus chrysogaster Moschus chrysogaster Moschus chrysogaster
Moschus cupreus Moschus cupreus Moschus cupreus Moschus cupreus
Moschus fuscus Moschus fuscus Moschus fuscus Moschus fuscus
Moschus leucogaster Moschus leucogaster Moschus leucogaster Moschus leucogaster
Moschus moschiferus Moschus moschiferus Moschus moschiferus Moschus moschiferus
 
Family Cervidae
Subfamily Capreolinae
Tribe Alceini
Alces
Alces alces
Alces americanus
Alces alces
Alces americanus
Alces alces Alces alces
 
Capreolus
Capreolus capreolus Capreolus capreolus Capreolus capreolus Capreolus capreolus
Capreolus pygargus Capreolus pygargus Capreolus pygargus Capreolus pygargus
Hydropotes
Hydropotes inermis Hydropotes inermis Hydropotes inermis Hydropotes inermis
 
Blastocerus
Blastocerus dichotomus Blastocerus dichotomus Blastocerus dichotomus Blastocerus dichotomus
Hippocamelus
Hippocamelus antisensis Hippocamelus antisensis Hippocamelus antisensis Hippocamelus antisensis
Hippocamelus bisulcus Hippocamelus bisulcus Hippocamelus bisulcus Hippocamelus bisulcus
Mazama
Mazama americana
Mazama bororo
Mazama americana
Mazama bororo
Mazama gualea
Mazama jucunda
Mazama trinitatis
Mazama whitelyi
Mazama zamora
Mazama zetta
Mazama americana
Mazama bororo
Mazama americana
Mazama jucunda
Mazama rufa
Mazama bricenii
Mazama rufina
Mazama bricenii
Mazama rufina
Mazama bricenii
Mazama rufina
Mazama rufina
Mazama chunyi Mazama chunyi Mazama chunyi Mazama chunyi
Mazama gouazoubira

Mazama cita
Mazama gouazoubira
Mazama murelia
Mazama nemorivaga
Mazama permira
Mazama sanctaemartae
Mazama superciliaris
Mazama tschudii
Mazama rondoni

Mazama gouazoubira
Mazama nemorivaga
Bisbalus citus
Passalites nemorivagus
Subulo gouazoubira
Mazama nana Mazama nana Mazama nana Mazama nanus
Mazama temama Mazama temama Mazama temama Mazama temama
Odocoileus
Odocoileus hemionus Odocoileus hemionus Odocoileus hemionus Odocoileus hemionus
Mazama pandora Mazama pandora Mazama pandora Odocoileus pandora
Odocoileus virginianus Odocoileus virginianus Odocoileus virginianus Odocoileus virginianus
(With a note that the genus Odocoileus requires significant taxonomic revision)
Ozotoceros
Ozotoceros bezoarticus Ozotoceros bezoarticus Ozotoceros bezoarticus Ozotoceros bezoarticus
Pudu
Pudu mephistophiles Pudu mephistophiles Pudu mephistophiles (Pudella carlae)
Pudella mephistophila
Pudu puda Pudu puda Pudu puda Pudu puda
Rangifer
Rangifer tarandus Rangifer tarandus Rangifer tarandus Rangifer tarandus
 
Subfamily Cervinae
Tribe Cervini
Axis
Axis axis Axis axis Axis axis Axis axis
Axis calamianensis Axis calamianensis Axis calamianensis Axis calamianensis
Axis kuhlii Axis kuhlii Axis kuhlii Axis kuhlii
Axis porcinus Axis annamiticus
Axis porcinus
Axis porcinus Axis porcinus
Cervus
Cervus elaphus Cervus alashanicus
Cervus bactrianus
Cervus canadensis
Cervus corsicanus
Cervus elaphus
Cervus hanglu
Cervus macneilli
Cervus maral
Cervus pannoniensis
Cervus wallichii
Cervus xanthopygus
Cervus yarkandensis
Cervus canadensis
Cervus elaphus
Cervus hanglu
Cervus canadensis
Cervus elaphus
Cervus hanglu
Cervus nippon Cervus aplodontus
Cervus hortulorum
Cervus nippon
Cervus pseudaxis
Cervus pulchellus
Cervus sichuanicus
Cervus taiouanus
Cervus nippon Cervus nippon
Przewalskium albirostris Cervus albirostris Cervus albirostris Cervus albirostris
Dama
Dama dama Dama dama
Dama mesopotamica
Dama dama
Dama mesopotamica
Dama dama
Dama mesopotamica
Elaphurus
Elaphurus davidianus Elaphurus davidianus Elaphurus davidianus Elaphurus davidianus
Rucervus
Rucervus duvaucelii Rucervus branderi
Rucervus duvaucelii
Rucervis ranjitsinhi
Rucervus duvaucelii Rucervus duvaucelii
Rucervus elsii Panolia eldii
Panolia siamensis
Panolia thamin
Rucervus eldii Rucervus eldii
Rucervus schomburgki Rucervus schomburgki Rucervus schomburgki Rucervus schomburgki
Rusa
Rusa alfredi Cervus alfredi Rusa alfredi Rusa alfredi
Rusa marianna Cervus barandanus
Cervus mariannus
Cervus nigellus
Rusa marianna Rusa marianna
Rusa timorensis Cervus timorensis Rusa timorensis Rusa timorensis
Rusa unicolor Cervus equinus
Cervus unicolor
Rusa unicolor Rusa unicolor
 
Elaphodus
Elaphodus cephalophus Elaphodus cephalophus Elaphodus cephalophus Elaphodus cephalophus
Muntiacus
Muntiacus atherodes Muntiacus atherodes Muntiacus atherodes Muntiacus atherodes
Muntiacus crinifrons Muntiacus crinifrons Muntiacus crinifrons Muntiacus crinifrons
Muntiacus feae Muntiacus feae Muntiacus feae Muntiacus feae
Muntiacus gongshanensis Muntiacus gongshanensis Muntiacus gongshanensis Muntiacus gongshanensis
Muntiacus muntjak

Muntiacus aureus
Muntiacus malabaricus
Muntiacus montanus
Muntiacus muntjak
Muntiacus nigripes
Muntiacus vaginalis

Muntiacus montanus
Muntiacus muntjak
Muntiacus vaginalis
Muntiacus malabaricus
Muntiacus muntjak
Muntiacus vaginalis
Muntiacus puhoatensis Muntiacus puhoatensis Muntiacus puhoatensis Muntiacus puhoatensis
Muntiacus putaoensis Muntiacus putaoensis Muntiacus putaoensis Muntiacus putaoensis
Muntiacus reevesi Muntiacus reevesi Muntiacus reevesi Muntiacus reevesi
Muntiacus rooseveltorum Muntiacus rooseveltorum Muntiacus rooseveltorum Muntiacus rooseveltorum
Muntiacus truongsonensis Muntiacus truongsonensis Muntiacus truongsonensis Muntiacus truongsonensis
Muntiacus vuquangensis Muntiacus vuquangensis Muntiacus vuquangensis Muntiacus vuquangensis
 
Family Bovidae
Subfamily Antilopinae
Tribe Aepycerotini
Aepyceros
Aepyceros melampus Aepyceros melampus
Aepyceros petersi
Aepyceros melampus Aepyceros melampus
 
Alcelaphus
Alcelaphus buselaphus
Alcelaphus caama
Alcelaphus lichtensteini
Alcelaphus buselaphus
Alcelaphus caama
Alcelaphus cokii
Alcelaphus lelwel
Alcelaphus lichtensteini
Alcelaphus major
Alcelaphus swaynei
Alcelaphus tora
Alcelaphus buselaphus Alcelaphus buselaphus
Beatragus
Beatragus hunteri Beatragus hunteri Beatragus hunteri Beatragus hunteri
Connochaetes
Connochaetes gnou Connochaetes gnou Connochaetes gnou Connochetes gnou
Connochaetes taurinus Connochaetes albojubatus
Connochaetes johnstoni
Connochaetes mearnsi
Connochaetes taurinus
Connochaetes taurinus Connochaetes taurinus
Damaliscus
Damaliscus korrigum
Damaliscus lunatus
Damaliscus superstes
Damaliscus eurus
Damaliscus jimela
Damaliscus korrigum
Damaliscus lunatus
Damaliscus selousi
Damaliscus superstes
Damaliscus tiang
Damaliscus topi
Damaliscus ugandae
Damaliscus lunatus Damaliscus lunatus
Damaliscus pygargus Damaliscus phillipsi
Damaliscus pygargus
Damaliscus pygargus Damaliscus pygargus
 
Tribe Antilopini
Ammodorcas
Ammodorcas clarkei Ammodorcas clarkei Ammodorcas clarkei Ammodorcas clarkei
Antidorcas
Antidorcas marsupialis Antidorcas angolensis
Antidorcas hofmeyri
Antidorcas marsupialis
Antidorcas marsupialis Antidorcas marsupialis
Antilope
Antilope cervicapra Antilope cervicapra Antilope cervicapra Antilope cervicapra
Dorcatragus
Dorcatragus megalotis Dorcatragus megalotis Dorcatragus megalotis Dorcatragus megalotis
Eudorcas
Eudorcas rufifrons Eudorcas albonotata
Eudorcas rufifrons
Eudorcas tilonura
Eudorcas albonotata
Eudorcas rufifrons
Eudorcas tilonura
Eudorcas albonotata
Eudorcas rufifrons
Eudorcas tilonura
Eudorcas rufina Eudorcas rufina  - Eudorcas rufina
Eudorcas thomsonii Eudorcas nasalis
Eudorcas thomsoni
Eudorcas thomsonii Eudorcas thomsonii
Gazella
Gazella arabica
Gazella erlangeri
Gazella acaciae
Gazella arabica
Gazella bilkis
Gazella cora
Gazella dareshurii
Gazella erlangeri
Gazella muscatensis
Gazella arabica
Gazella bilkis
Gazella arabica
Gazella bennettii Gazella bennettii
Gazella christyi
Gazella fuscifrons
Gazella shikarii
Gazella salinarum
Gazella bennettii Gazella bennettii
Gazella cuvieri Gazella cuvieri Gazella cuvieri Gazella cuvieri
Gazella dorcas
Gazella saudiya
Gazella dorcas
Gazella pelzelni
Gazella saudiya
Gazella dorcas
Gazella saudiya
Gazella dorcas

Gazella gazella

Gazella gazella
Gazella karamii
Gazella gazella Gazella gazella
Gazella leptoceros Gazella leptoceros Gazella leptoceros Gazella leptoceros
Gazella spekei Gazella spekei Gazella spekei Gazella spekei
Gazella subgutturosa Gazella gracilicornis
Gazella marica
Gazella subgutturosa
Gazella yarkandensis
Gazella marica
Gazella subgutturosa
Gazella marica
Gazella subgutturosa
Litocranius
Litocranius walleri Litocranius sclateri
Litocranius walleri
Litocranius walleri Litocranius walleri
Madoqua
Madoqua guentheri Madoqua guentheri
Madoqua smithii
Madoqua guentheri Madoqua guentheri
Madoqua kirkii Madoqua cavendishi
Madoqua damarensis
Madoqua hindei
Madoqua kirkii
Madoqua thomasi
Madoqua kirkii Madoqua cavendishi
Madoqua damarensis
Madoqua kirkii
Madoqua thomasi
Madoqua piacentinii Madoqua piacentinii Madoqua piacentinii Madoqua piacentinii
Madoqua saltiana Madoqua hararensis
Madoqua lawrancei
Madoqua phillipsi
Madoqua saltiana
Madoqua swaynei
Madoqua saltiana Madoqua saltiana
Nanger
Nanger dama Nanger dama Nanger dama Nanger dama
Nanger granti Nanger granti
Nanger notata
Nanger petersi
Nanger granti Nanger granti
Nanger notatus
Nanger petersii
Nanger soemmerringii Nanger soemmerringi Nanger soemmerringii Nanger soemmerringii
Ourebia
Ourebia ourebi Ourebia hastata
Ourebia montana
Ourebia ourebi
Ourebia quadriscopa
Ourebia ourebi Ourebia ourebi
Procapra
Procapra gutturosa Procapra gutturosa Procapra gutturosa Procapra gutturosa
Procapra picticaudata Procapra picticaudata Procapra picticaudata Procapra picticaudata
Procapra przewalskii Procapra przewalskii Procapra przewalskii Procapra przewalskii
Raphicerus
Raphicerus campestris Raphicerus campestris Raphicerus campestris Raphicerus campestris
Raphicerus melanotis Raphicerus melanotis Raphicerus melanotis Raphicerus melanotis
Raphicerus sharpei Raphicerus colonicus
Raphicerus sharpei
Raphicerus sharpei Raphicerus sharpei
Saiga
Saiga tatarica Saiga mongolica
Saiga tartarica
Saiga tatarica Saiga tatarica
 
Tribe Caprini
Ammotragus
Ammotragus lervia Ammotragus lervia Ammotragus lervia Ammotragus lervia
Arabitragus
Hemitragus jayakari Arabitragus jayakari Arabitragus jayakari Arabitragus jayakari
Budorcas
Budrocas taxicolor Budorcas bedfordi
Budorcas taxicolor
Budorcas tibetana
Budorcas whitei
Budorcas taxicolor Budorcas taxicolor
Budorcas tibetana
Capra
Capra caucasica Capra caucasica
Capra cylindricornis
Capra caucasica
Capra cylindricornis
Capra caucasica
Capra cylindricornis
Capra falconeri Capra falconeri Capra falconeri Capra falconeri
Capra hircus Capra aegagrus Capra aegagrus Capra aegagrus
Capra hircus
Capra ibex Capra ibex Capra ibex Capra ibex
Capra nubiana Capra nubiana Capra nubiana Capra nubiana
Capra pyrenaica Capra pyrenaica Capra pyrenaica Capra pyrenaica
Capra sibirica Capra sibirica Capra sibirica Capra sibirica
Capra walie Capra walie Capra walie Capra walie
Capricornis
Capricornis crispus Capricornis crispus Capricornis crispus Capricornis crispus
Capricornis milneedwardsi
Capricornis sumatrensis
Capricornis thar

Capricornis maritimus
Capricornis milneedwardsi
Capricornis sumatrensis
Capricornis thar

Capricornis sumatrensis Capricornis sumatrensis
Capricornis rubidus Capricornis rubidus Capricornis rubidus Capricornis rubidus
Capricornis swinhoei Capricornis swinhoii Capricornis swinhoei Capricornis swinhoei
Hemitragus
Hemitragus jemlahicus Hemitragus jemlahicus Hemitragus jemlahicus Hemitragus jemlahicus
Naemorhedus
Naemorhedus baileyi Nemorhaedus baileyi Naemorhedus baileyi Naemorhedus baileyi
Naemorhedus cranbrooki
Naemorhedus caudatus Nemorhaedus caudatus Naemorhedus caudatus Naemorhedus caudatus
Naemorhedus goral Nemorhaedus bedfordi
Nemorhaedus goral
Naemorhedus goral Naemorhedus goral
Naemorhedus griseus Nemorhaedus evansi
Nemorhaedus griseus
(Naemorhedus griseus) Naemorhedus evansi
Naemorhedus griseus
Nilgiritragus
Nilgiritragus hylocrius Nilgiritragus hylocrius Nilgiritragus hylocrius Nilgiritragus hylocrius
Oreamnos
Oreamnos americanus Oreamnos americanus Oreamnos americanus Oreamnos americanus
Ovibos
Ovibos moschatus Ovibos moschatus Ovibos moschatus Ovibos moschatus
Ovis
Ovis ammon Ovis ammon
Ovis collium
Ovis darwini
Ovis hodgsoni
Ovis jubata
Ovis karelini
Ovis nigrimontana
Ovis polii
Ovis ammon Ovis ammon
Ovis aries Ovis arabica
Ovis bochariensis
Ovis cycloceros
Ovis gmelini
Ovis isphaganica
Ovis laristanica
Ovis punjabensis
Ovis severtzovi
Ovis vignei
Ovis gmelini
Ovis vignei
Ovis aries
Ovis gmelinii
Ovis vignei
Ovis canadensis Ovis canadensis Ovis canadensis Ovis canadensis
Ovis dalli Ovis dalli Ovis dalli Ovis dalli
Ovis nivicola Ovis nivicola Ovis nivicola Ovis nivicola
Pantholops
Pantholops hodgsonii Pantholops hodgsonii Pantholops hodgsonii Pantholops hodgsonii
Pseudois
Pseudois nayaur
Pseudois schaeferi
Pseudois nayaur
Pseudois schaeferi
Pseudois nayaur Pseudois nayaur
Rupicapra
Rupicapra pyrenaica Rupicapra ornata
Rupicapra parva
Rupicapra pyrenaica
Rupicapra pyrenaica Rupicapra pyrenaica
Rupicapra rupicapra Rupicapra asiatica
Rupicapra carpatica
Rupicapra rupicapra
?Rupicapra balcanica
Rupicapra rupicapra Rupicapra rupicapra
 
Cephalophorus
Cephalophus brookei
Cephalophus ogilbyi
Cephalophus brookei
Cephalophus crusalbum
Cephalophus ogilbyi
Cephalophus ogilbyi Cephalophorus brookei
Cephalophorus ogilbyi
Cephalophus callipygus Cephalophus callipygus Cephalophus callipygus Cephalophorus callipygus
Cephalophus leucogaster Cephalophus arrhenii
Cephalophus leucogaster
Cephalophus leucogaster Cephalophorus leucogaster
Cephalophus natalensis Cephalophus harveyi
Cephalophus natalensis
Cephalophus harveyi
Cephalophus natalensis
Cephalophorus natalensis
Cephalophus niger Cephalophus niger Cephalophus niger Cephalophorus niger
Cephalophus nigrifrons Cephalophus fosteri
Cephalophus hooki
Cephalophus hypoxanthus
Cephalophus nigrifrons
Cephalophus rubidus
Cephalophus nigrifrons Cephalophorus kivuensis
Cephalophorus nigrifrons
Cephalophorus rubidus
Cephalophus rufilatus Cephalophus rufilatus Cephalophus rufilatus Cephalophorus rufilatus
Cephalophus weynsi Cephalophus johnstoni
Cephalophus lestradei
Cephalophus weynsi
Cephalophus weynsi Cephalophorus weynsi
Cephalophula
Cephalophus zebra Cephalophus zebra Cephalophus zebra Cephalophula zebra
Cephalophus
Cephalophus dorsalis Cephalophus castaneus
Cephalophus dorsalis
Cephalophus dorsalis Cephalophus dorsalis
Cephalophus jentinki Cephalophus jentinki Cephalophus jentinki Cephalophus jentinki
Cephalophus silvicultor Cephalophus curticeps
Cephalophus silvicultor
Cephalophus silvicultor Cephalophus silvicultor
Cephalophus spadix Cephalophus spadix Cephalophus spadix Cephalophus spadix
Leucocephalophus
Cephalophus adersi Cephalophus adersi Cephalophus adersi Leucocephalophus adersi
Philantomba
Philantomba maxwellii Philantomba maxwelli
Philantomba walteri
Philantomba maxwellii
Philantomba walteri
Philantomba maxwellii
Philantomba walteri
Philantomba monticola Philantomba aequatorialis
Philantomba anchietae
Philantomba bicolor
Philantomba congicus
Philantomba defriesi
Philantomba hecki
Philantomba lugens
Philantomba melanorheus
Philantomba monticola
Philantomba simpsoni
Philantomba monticola Philantomba monticola
Sylvicapra
Sylvicapra grimmia Sylvicapra coronatus
Sylvicapra grimmia
Sylvicapra pallidior
Sylvicapra grimmia Sylvicapra grimmia
 
Tribe Hippotragini
Addax
Addax nasomaculatus Addax nasomaculatus Addax nasomaculatus Addax nasomaculatus
Hippotragus
Hippotragus equinus Hippotragus equinus Hippotragus equinus Hippotragus equinus
Hippotragus leucophaeus Hippotragus leucophaeus Hippotragus leucophaeus Hippotragus leucophaeus
Hippotragus niger Hippotragus niger
Hippotragus roosevelti
Hippotragus niger Hippotragus niger
Oryx
Oryx beisa Oryx beisa
Oryx callotis
Oryx gallarum
Oryx beisa Oryx beisa
Oryx dammah Oryx dammah Oryx dammah Oryx dammah
Oryx gazella Oryx gazella Oryx gazella Oryx gazella
Oryx leucoryx Oryx leucoryx Oryx leucoryx Oryx leucoryx
 
Tribe Neotragini
Neotragus
Neotragus pygmaeus Neotragus pygmaeus Neotragus pygmaeus Neotragus pygmaeus
Nesotragus
Neotragus batesi Neotragus batesi Neotragus batesi Nesotragus batesi
Neotragus moschatus

Neotragus kirchenpaueri Neotragus livingstonianus
Neotragus moschatus

Nesotragus moschatus Nesotragus moschatus
 
Tribe Oreotragini
Oreotragus
Oreotragus oreotragus Oreotragus aceratos
Oreotragus aureus
Oreotragus centralis
Oreotragus oreotragus
Oreotragus porteousi
Oreotragus saltatrixoides
Oreotragus schillingsi
Oreotragus somalicus
Oreotragus stevensoni
Oreotragus transvaalensis
Oreotragus tyleri
Oreotragus oreotragus Oreotragus oreotragus
 
Tribe Reduncini
Kobus
Kobus ellipsiprymnus Kobus defassa
Kobus ellipsiprymnus
Kobus ellipsiprymnus Kobus ellipsiprymnus
Kobus kob Kobus kob
Kobus leucotis
Kobus loderi
Kobus thomasi
Kobus kob Kobus kob
Kobus leche Kobus anselli
Kobus kafuensis
Kobus leche
Kobus robertsi
Kobus smithemani
Kobus leche Kobus leche
Kobus megaceros Kobus megaceros Kobus megaceros Kobus megaceros
Kobus vardonii Kobus vardoni Kobus vardonii Kobus vardonii
Pelea
Pelea capreolus Pelea capreolus Pelea capreolus Pelea capreolus
Redunca
Redunca arundinum Redunca arundinum
Redunca occidentalis
Redunca arundinum Redunca arundinum
Redunca fulvorufula Redunca adamauae
Redunca chanleri
Redunca fulvorufula
Redunca fulvorufula Redunca fulvorufula
Redunca redunca Redunca bohor
Redunca cottoni
Redunca nigeriensis
Redunca redunca
Redunca redunca Redunca redunca
 
Subfamily Bovinae
Tribe Boselaphini
Boselaphus
Boselaphus tragocamelus Boselaphus tragocamelus Boselaphus tragocamelus Boselaphus tragocamelus
Tetracerus
Tetracerus quadricornis Tetracerus quadricornis Tetracerus quadricornis Tetracerus quadricornis
 
Tribe Bovini
Bos
Bison bison Bos bison Bison bison Bos bison
Bison bonasus Bos bonasus
Bos caucasicus
Bison bonasus Bos bonasus
Bos frontalis Bos gaurus Bos gaurus Bos frontalis
Bos gaurus
Bos grunniens Bos mutus Bos mutus Bos grunniens
Bos mutus
Bos javanicus Bos javanicus Bos javanicus Bos javanicus
Bos sauveli Bos sauveli Bos sauveli Bos sauveli
Bos taurus Bos primigenius Bos primigenius Bos primigenius
Bos taurus
Bubalus
Bubalus bubalis Bubalus arnee Bubalus arnee Bubalus arnee
Bubalus bubalis
Bubalus depressicornis Bubalus depressicornis Bubalus depressicornis Bubalus depressicornis
Bubalus mindorensis Bubalus mindorensis Bubalus mindorensis Bubalus mindorensis
Bubalus quarlesi Bubalus quarlesi Bubalus quarlesi Bubalus quarlesi
Pseudoryx
Pseudoryx nghetinhensis Pseudoryx nghetinhensis Pseudoryx nghetinhensis Pseudoryx nghetinhensis
Syncerus
Syncerus caffer Syncerus brachyceros
Syncerus caffer
Syncerus mathewsi
Syncerus nanus
Syncerus caffer Syncerus caffer
 
Tribe Tragelaphini
Tragelaphus
Taurotragus derbianus Taurotragus derbianus Tragelaphus derbianus Tragelaphus derbianus
Taurotragus oryx Taurotragus oryx Tragelaphus oryx Tragelaphus oryx
Tragelaphus angasii Nyala angasii Tragelaphus angasii Tragelaphus angasii
Tragelaphus buxtoni Tragelaphus buxtoni Tragelaphus buxtoni Tragelaphus buxtoni
Tragelaphus eurycerus Tragelaphus euryceros Tragelaphus eurycerus Tragelaphus eurycerus
Tragelaphus imberbis Ammelaphus australis
Ammelaphus imberbis
Tragelaphus imberbis Tragelaphus imberbis
Tragelaphus scriptus Tragelaphus bor
Tragelaphus decula
Tragelaphus fasciatus
Tragelaphus meneliki
Tragelaphus ornatus
Tragelaphus phaleratus
Tragelaphus scriptus
Tragelaphus sylvaticus
Tragelaphus scriptus Tragelaphus scriptus
Tragelaphus sylvaticus
Tragelaphus spekii Tragelaphus gratus
Tragelaphus larkenii
Tragelaphus selousi
Tragelaphus spekii
Tragelaphus sylvestris
?Tragelaphus ugallae
Tragelaphus spekii Tragelaphus spekii
Tragelaphus strepsiceros Strepsiceros chora
Strepsiceros cottoni
Strepsiceros strepsiceros
Strepsiceros zambesiensis
Tragelaphus strepsiceros Tragelaphus strepsiceros

Literature Cited

American Society of Mammalogists (2024). Mammal Diversity Database v1.13. Available online at the Mammal Diversity Database website.

Groves, C., and Grubb, P. (2011). Ungulate Taxonomy. The John Hopkins University Press, Baltimore.

IUCN (International Union for Conservation of Nature and Natural Resources) (2024). IUCN Red List of Threatened Species. Version 2023-1. Available online at the IUCN Redlist website.

Wilson, D. E., and D. M. Reeder [editors]. 2005.  Mammal Species of the World (3rd Edition).  Johns Hopkins University Press, 2,142 pp. Available online at the Mammal Species of the World website.